The Center is the player most responsible for defense. He or she typically lines up at the center hash mark at the front of the team area, though there are various defensive arrangements. The center is the most defensive position simply because he or she must be able to move both to the left and right to defend. He or she is also typically the defensive coordinator, as he or she can more accurately determine which opposing player has the ball due to being square to the opposing team.
The Left and Right Wings generally line up at the end of the hash marks coming off of the respective sidelines. Their main defensive responsibilities are keeping the ball out of either corner of the goal, though they also defend some towards the center. Typically, the wings are the main offense, allowing the Center to reserve his or her energy for defense, while they attempt to score. This is not to say that the Center is not also an offensive threat, just that he or she generally is more focused on keeping the ball out of his or her own net than putting it into the opponents.